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Ripple W.J., Larsen E.J., Renkin R.A., Smith D.W.. 2001. Trophic Cascades among wolves, elk and aspen on Yellowstone National Park's northern range.. Biological Conservation. 102:227-234.

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Ripple W.J., Estes J.A., Beschta R.L., Wilmers C.C., Ritchie E.G., Hebblewhite M., Berger J., Elmhagen B., Letnic M., Nelson M.P et al.. 2014. Status and Ecological Effects of the World's Largest Carnivores. Science. 343(6167)